Do you get the same problem using the actual dcarter-businessone skin? I do not.
All I can guess is you didn't copy all the files or you modified them, particularly the theme.skin and layout.master file.
Else if you are running in .NET 3.5 under Medium Trust hosting then it can't use the theme.skin file from the skin folder, it can only use the one from /App_Themes/default, so you would need to copy the theme.skin file from the skin folder into the /App_Themes/default folder. This is discussed at the bottom of our article Understanding the theme.skin file.
Hope that helps,